Borneo

Muludeercaveimgp5935 438x0 scale — Limestone dipterocarp lowland rainforest, Deer Cave, Mulu National Park, Sarawak, Borneo. @ C.Michael Hogan [1]

This is a checklist of Opiliones species of the island of Borneo (including Kalimantan, Sarawak, Sabah & Brunei).

Suborder Cyphophthalmi

Family Stylocellidae
- - Miopsalis collinsi (Shear, 1993a):181
  - Miopsalis gryllospeca (Shear, 1993a):179
  - Miopsalis kinabalu (Shear, 1993a):184
  - Miopsalis leakeyi (Shear, 1993a):185
  - Miopsalis lionota (Pocock, 1897):291
  - Miopsalis mulu (Shear, 1993a):182
  - Miopsalis pocockii (Hansen & Sørensen, 1904):97
  - Miopsalis sabah (Shear, 1993a):185
  - Miopsalis silhavyi (Rambla, 1991):228

Suborder Dyspnoi (none)

Suborder Eupnoi

Family Sclerosomatidae
- Subfamily Gagrellinae
  - Bastia guttata Banks, 1930a:77
  - Chasenella luma Roewer, 1933b:46
  - Chasenella pakka Roewer, 1933b:47
  - Dentobunus bicorniger (Simon, 1901):81
  - Dentobunus bidentatus (Thorell, 1891c):697
  - Dentobunus luteus Roewer, 1910a:39
  - Dentobunus pulcer Suzuki, 1969b::72
  - Eugagrella paupera Banks, 1930a:81 - See Roewer, 1954c:287
  - Eugagrella paupera apicalis Banks, 1930a:81
  - Euzaleptus sarawakensis Roewer, 1911b:182
  - Gagrella borneoensis Roewer, 1954b:216
  - Gagrella corrugata Roewer, 1954b:217
  - Gagrella infuscata Roewer, 1911a:252
  - Gagrella insculpta Pocock, 1897:283
  - Gagrella lateritia Roewer, 1954b:217
  - Gagrella longipalpis Thorell, 1891c:693
  - Gagrella mjobergi (Banks, 1930a):79
  - Gagrella paupera With, 1905:7
  - Gagrella pretiosa (Banks, 1930a):80
  - Gagrella rorida Roewer, 1954b:216
  - Gagrella sampitia Roewer, 1954b:216
  - Gagrella sarawakensis With, 1905:5
  - Gagrella scorbiculata Thorell, 1891c:717
  - Gagrella thorelli (Banks, 1930a):79
  - Gagrellula curvispina Roewer, 1912b:45
  - Hologagrella curvispina Banks, 1930a:81
  - Hologagrella normalis Banks, 1930a:81
  - Marthana affinis Banks, 1930a:85
  - Marthana columnaris Thorell, 1891c:722
  - Marthana furcata Banks, 1930a:85
  - Marthana vestita With, 1905:9
  - Melanopa cinctipes (Banks, 1930a):83
  - Melanopa maculipes (Banks, 1930a):83
  - Melanopa nigripes (Banks, 1930a0):84
  - Metazaleptus borneensis Banks, 1930a:75
  - Metazaleptus montanus Banks, 1930a:74
  - Metazaleptus palpalis Banks, 1930a:75
  - Metazaleptus rufescens Banks, 1930a:76
  - Paragagrella brevispina Banks, 1930a:82
  - Psathyropus granulata (Roewer, 1954c):246
  - Toragrella longipes Roewer, 1955a:74
  - Zaleptiolus laevipes Roewer, 1955b:169
  - Zaleptulus unicolor Roewer, 1955b:166
  - Zaleptulus banksi Roewer, 1955b:168
  - Zaleptulus lineatus Roewer, 1955b:168
  - Zaleptus trichopus Thorell, 1876b:116
  - Zaleptus annulipes Banks, 1930a:73
  - Zaleptus lugubris (Thorell, 1889):621
  - Zaleptus mjobergi (Banks, 1930a):76
  - Zaleptus pulchellus Banks, 1930a:73
  - Zaleptus quadricornis (Thorell, 1891c):689

Suborder Laniatores (Grassatores)

- Grassatores incertae sedis
  - Arulla parvula Suzuki, 1969b:20
  - Istithaeus atroluteus Roewer, 1949a:49
  - Kondosus borneoensis Roewer, 1949a:49
Family Assamiidae
- Subfamily Dampetrinae
  - Dunkeriana borneensis Roewer, 1935a:127
  - Mermerus thorelli Banks, 1930a:69
  - Metahyamus amoensis Suzuki, 1969b:49
  - Metahyamus bicolor Roewer, 1935a:132
- Subfamily Trionyxellinae
  - Dulitellus maculatus Roewer, 1935a:147
  - Dulitellus opacus (Roewer, 1935a):147
  - Dulitellus sarawakensis Suzuki, 1969b:46
Family Epedanidae
- Subfamily (undetermined) Epedanidae incertae sedis
  - Parabupares robustus Suzuki, 1982d:100
  - Sotekia minima Suzuki, 1982d:103
- Subfamily Epedaninae
  - Epedanus brevipalpus Banks, 1930a:60
  - Epedanus cavicolus Banks, 1930a:60
  - Epedanus lutescens Thorell, 1876b:133
  - Epedanus pictus Thorell, 1876b:128
  - Epedanus praedo Sørensen, 1932:254
  - Heteroepedanus monacantha (Roewer, 1911d):601
  - Heteroepedanus triacantha (Roewer, 1911d):603
  - Lobonychium palpiplus Roewer, 1938b:125
  - Metepedanulus flaveolus Banks, 1930a:62
  - Metepedanus venator (Roewer, 1911b):184
  - Metepedanus accentuatus (Roewer, 1911b):186
  - Nanepedanus rufus Roewer, 1938b:103
- Subfamily Acrobuninae
  - Acrobunus thorelli Banks, 1930a:61
  - Metacrobunus frontalis Banks, 1930a:65
- Subfamily Sarasinicinae
  - Albertops robustus Roewer, 1938b:157
  - Asopella lutescens Thorell, 1876b:133
  - Koyanus clarus Roewer, 1938b:163
  - Kuchingius megalopalpus Roewer, 1927a:333
  - Opelytus rugichelis Roewer, 1938b:150
  - Punanus tenuis Roewer, 1938b:142
  - Sarasinica atra Roewer, 1938b:134
  - Sarasinica femoralis Roewer, 1938b:134
  - Sarasinica tricomma Roewer, 1914a:90
  - Sinistus maculatus Roewer, 1938b:152
  - Sinistus fuscus Roewer, 1938b:152
  - Tegestria borneensis Roewer, 1938b:145
Family Podoctidae
- Subfamily Ibaloniinae
  - Bonea albertus (Roewer, 1949c):258
  - Bonea armatissima (Roewer, 1949c):262
  - Bonea silvestris (Roewer, 1949c):260
  - Metibalonius oppositus (Roewer, 1927a):308
- Subfamily Podoctinae
  - Baramella quadrispina (Roewer, 1915c):52
  - Baramia echinosa Banks, 1930a:63
  - Baramia longipes Banks, 1930a:64
  - Baramia vorax Hirst, 1912:76
  - Gaditusa coxalis Roewer, 1949c:277
  - Japetus longipes Roewer, 1949c:274
  - Lejokus silvestris Roewer, 1949c:272
  - Lundulla bifurcata Roewer, 1927a:319
  - Trencona setipes Roewer, 1949c:277
  - Trigonobunus spinifer Loman, 1894:154
Family Sandokanidae
Gnomulus annulipes (Pocock, 1897):283
- - Gnomulus baharu Schwendinger, 1998:527
  - Gnomulus carinatus Schwendinger & Martens, 2002a:87
  - Gnomulus conigerus (Schwendinger 1992):183
  - Gnomulus exsudans Schwendinger & Martens, 2002a:83
  - Gnomulus hutan Schwendinger & Martens, 2002a:81
  - Gnomulus imadatei (Suzuki, 1969b):12
  - Gnomulus laevis (Roewer, 1915c):128
  - Gnomulus lomani Schwendinger & Martens, 2002a:77
  - Gnomulus obscurus Schwendinger & Martens, 2002a:80
  - Gnomulus sundaicus (Schwendinger 1992):185
  - Martensiellus tenuipalpus Schwendinger, 2006:258
  - Sandokan doriae (Thorell, 1876b):135
  - Sandokan expatriatus (Schwendinger & Martens, 2004):167 {More likely than Thailand - see note on species page}
  - Sandokan hosei (Pocock, 1897):285
  - Sandokan kaltim (Schwendinger, 2007):729
  - Sandokan megachelis (Schwendinger, 1992):192
Family Tithaeidae
- - Metatithaeus rubidus Suzuki, 1969b:29
  - Tithaeus borneensis Roewer, 1949a:45
  - Tithaeus flavescens Banks, 1930a:68
  - Tithaeus fuscus Roewer, 1949a:45
  - Tithaeus granulatus Banks, 1930a:69
  - Tithaeus lesserti Roewer, 1949:45
  - Tithaeus longipes Banks, 1930a:67
  - Tithaeus nigripes Banks, 1930a:67
  - Tithaeus pumilio Roewer, 1949a:46
  - Tithaeus rotundus Suzuki, 1969b:26
  - Tithaeus sarawakensis Roewer, 1912c:121

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Literature[]

Banks, N. (1930a) Phalangida from Borneo. The Sarawak Museum Journal, n.s., Vol. 4 (Part I), No. 12, pp. 57–86
Martens, J. & Schwendinger, P. (1998) A taxonomic revision of the family Oncopodidae I. New genera and new species of Gnomulus Thorell (Opiliones, Laniatores). Revue suisse de Zoologie, 105(3), 499–555.
Pocock, R.I. (1897) Description of some new Oriental Opiliones recently received by the British Museum. The Annals and Magazine of Natural History, London, (series 6) 19(111), 283–292.
Rambla, M. (1991) A new Stylocellus from caves of Borneo, Malaysia (Opiliones, Cyphophthalmi, Stylocellidae). Mémoires de Biospéologie, Moulis, 18, 227–232.
Roewer, C.F. (1910a) Revision der Opiliones Plagiostethi (= Opiliones Palpatores). I. Teil: Familie der Phalangiidae. (Subfamilien: Gagrellini, Liobunini, Leptobunini.) Abhandlungen aus dem Gebiete der Naturwissenschaften, herausgegeben vom Naturwissenschaftlichen Verein in Hamburg, Hamburg, 19(4), 1–294.
Roewer, C.F. (1911a) Note XX. Niederländisch-Indische Opiliones (Gagrellini) des Leidener Museums. Notes from the Leyden Museum, Leyden, 33, 249–260.
Roewer, C.F. (1911b) Opiliones aus Britisch Indien und Sarawak. Archiv für Naturgeschichte, Berlin, Abt. A, Original-Arbeiten, 77(2), 160-188.
Roewer, C.F. (1911d) Ost-asiatische Opiliones. Zoologische Jahrbücher, Jena, Abteilung für Systematik, Ökologie und Geographie der Tiere, 31(5), 591–612.
Roewer, C.F. (1912b) Einige neue Gattungen und Arten der Opiliones Palpatores aus den Subfamilien der Gagrellinae und Liobuninae der Familie der Phalangiidae. Archiv für Naturgeschichte, Berlin, Abt. A, Original-Arbeiten, 78(1), 27–59.
Roewer, C.F. (1912c) Die Familien der Assamiiden und Phalangodiden der Opiliones-Laniatores. (= Assamiden, Dampetriden, Phalangodiden, Epedaniden, Biantiden, Zalmoxiden, Samoiden, Palpipediden anderer Autoren). Archiv für Naturgeschichte, Berlin, Abt. A, Original-Arbeiten, 78(3), 1–242.
Roewer, C.F. (1914a) Die Opiliones der Sammlung der Herren Drs. Paul u. Fritz Sarasin auf Celebes in den Jahren 1893–1896. Archiv für Naturgeschichte, Berlin, Abt. A, Original-Arbeiten, 79(10) [1913], 70-96.
Roewer, C.F. (1915c) 106 neue Opilioniden. Archiv für Naturgeschichte, Berlin, Abt. A, Original-Arbeiten, 81(3), 1–152.
Roewer, C.F. (1933b) Die Opilioniden des Mt. Kinabalu, Nord-Borneo, 13,455 ft. Bulletin of the Raffles Museum, Singapore, 8, 46–47.
Roewer, C.F. (1935a) Alte und neue Assamiidae. Weitere Weberknechte VIII (8. Ergänzung der "Weberknechte der Erde" 1923). Veröffentlichungen aus dem Deutschen Kolonial- und Übersee-Museum in Bremen, Bremen, 1(1), 1–168.
Roewer, C.F. (1938b) Über Acrobuninae, Epedaninae und Sarasinicinae. Weitere Weberknechte IX. (9. Erganzung der "Weberknechte der Erde" 1923). Veröffentlichungen aus dem Deutschen Kolonial- und Übersee-Museum in Bremen, Bremen, 2(2), 81–169.
Roewer, C.F. (1949a) Über Phalangodiden I. (Subfam. Phalangodinae, Tricommatinae, Samoinae.) Weitere Weberknechte XIII. Senckenbergiana, 30(1/3), 11–61.
Roewer, C.F. (1949c) Über Phalangodidae II. Weitere Weberknechte XIV. Senckenbergiana, 30(4/6), 247–289.
Roewer, C.F. (1954b) Indoaustralische Gagrellinae (Opiliones, Arachnidae). (Weitere Weberknechte XVIII). 1. Teil. Senckenbergiana Biologica, Frankfurt, 35(3/4), 181–236.
Roewer, C.F. (1954c) Indoaustralische Gagrellinae (Opiliones, Arachnidae). (Weitere Weberknechte XVIII). 2. Teil. Senckenbergiana Biologica, Frankfurt, 35(5/6), 237–292.
Roewer, C.F. (1955a) Indoaustralische Gagrellinae (Opiliones, Arachnidae). (Weitere Weberknechte XVIII). 3. Teil. Senckenbergiana Biologica, Frankfurt, 36(1/2), 71–121.
Roewer, C.F. (1955b) Indoaustralische Gagrellinae (Opiliones, Arachnidae). (Weitere Weberknechte XVIII). 4. Teil (Schluss). Senckenbergiana Biologica, Frankfurt, 36(3/4), 123–171.
Schwendinger, P. (1992) New Oncopodidae (Opiliones, Laniatores) from Southeast Asia. Revue suisse de zoologie, Genève, 99(1), 177–199.
Schwendinger, P.J. (2006) A taxonomic revision of the family Oncopodidae VI. Martensiellus, a new genus from Borneo, and the discovery of a tarsal pore organ in Oncopodidae (Opiliones: Laniatores).Zootaxa, 1325, 255–266.
Schwendinger, P.J. (2007) A taxonomic revision of the family Oncopodidae VII. A new Oncopus species (Opiliones, Laniatores) from eastern Kalimantan. Revue Suisse de Zoologie, 114(4), 729–733.
Schwendinger, P.J. & Martens, J. (2002a) A taxonomic revision of the family Oncopodidae III. Further new species of Gnomulus Thorell (Opiliones, Laniatores). Revue suisse de zoologie, Genève, 109 (1), 47–113.
Schwendinger, P.J. & Martens, J. (2004) A taxonomic revision of the family Oncopodidae IV. The genus Oncopus Thorell (Opiliones, Laniatores). Revue suisse de zoologie, Genève, 111(1), 139–174.
Shear, W.A. (1993a) New species in the opilionid genus Stylocellus from Malaysia, Indonesia and the Philippines (Opiliones, Cyphophthalmi, Stylocellidae). Bulletin of the British Arachnological Society, 9, 174–188.
Sørensen, W.E. (1932) Descriptiones Laniatorum (Arachnidorum Opilionum Subordinis). Opus posthumum recognovit et edidit Kai L. Henriksen. Det Kongelige Danske Videnskabernes Selskabs skrifter [= Mémoires de l'Académie Royale des Sciences et des Lettres de Danemark], København [Copenhague], Naturvidenskabelig og Mathematisk Afdeling [= Section des sciences Naturelles et mathematiques], ser. 9, 3(4), 197–422.
Suzuki, S. (1969b) On a collection of opilionids from Southeast Asia. Journal of Science of the Hiroshima University, Series B, Division 1 (Zoology), Hiroshima, 22(2), 11–77.
Suzuki, S. (1982d) Two new genera of Phalangodinae (Opiliones, Phalangodidae) from Eastern Kalimantan, Borneo. Annotationes Zoologicae Japonenses, 55(2), 100-104.
Thorell, T.T.T. (1876b) Descrizione di alcune specie di Opilioni dell' Arcipelago Malese appartenenti al Museo Civico di Genova. Annali del Museo Civico di Storia Naturale di Genova, Genova, (series 1) 9 [“1876-1877”], 111–138.
Thorell, T.T.T. (1889) Aracnidi Artrogastri Birmani raccolti da L. Fea nel 1885–1887. In: Viaggio di Leonardo Fea in Birmania e regione vicine. XXI. Annali del Museo Civico di Storia Naturale di Genova, Genova, series 2, 7(3), 521–729.
Thorell, T.T.T. (1891c) Opilioni nuovi o poco conosciuti dell´Arcipelago Malese. Annali del Museo Civico di Storia Naturale di Genova, Genova, series 2, 10 (= 30) [“1890-1891”], 669–770.
With, C. J. (1905) Remarks on the Gagrellinae Thor., a group of Opiliones, with descriptions of some new species from Borneo. Bollettino dei Musei di Zoologia e di Anatomia Comparata della Reale Università di Torino, Torino, 20(509), 1–12.

Borneo

Literature[]

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